PALAEONTOS 17 ISSN: 1377-4654
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Landau B., Marques da Silva C. & Gili C., 2009, The Early Pliocene
Gastropoda (Mollusca) of Estepona, Southern Spain. Part 8: Nassariidae.
101 textpages, 3 textfigures, 1 table, 22 plates
Summary: In this
part of the series, giving a systematic account of the Gastropoda of
the Lower Pliocene (Zanclean) deposits of Estepona, province of
Málaga, Spain, the Nassariidae are described and discussed. The
study area has been expanded to include other major Iberian Pliocene
deposits, those of the Mondego basin of central-west Portugal, the
Guadalquivir basin of southern Atlantic Spain and the Alt
Empordà (Girona), Baix Llobregat (Barcelona) and Baix Ebre
(Tarragona), all northern Mediterranean Spain. The faunal composition
is compared with that present in the Atlantic, North Sea Basin and
Mediterranean Miocene to Recent times. The diversity of Nassariidae in
the Iberian assemblages is comparable in number of species and
composition to that found in the Italian Pliocene. 60 species are
described and discussed, with particular emphasis on protoconch
morphology. Ten species are new to science, Nassarius
iberoclathratus nov. sp., Nassarius patnuttalli nov nom., Nassarius
dinizi nov. sp., Nassarius plioholsaticus nov. sp., Nassarius
manuelmolinai nov. sp., Nassarius cotteri nov. sp., Nassarius
lusitanicus nov. sp., Nassarius mondegoensis nov. sp., Demoulia
zbyszewskii nov. sp. and Cyllenina amaldiformis nov.
sp. One further species we consider new, Nassarius sp. indet.
an nov sp., but do not name formally due to lack of well-preserved
specimens. Several new subjective synonymies are suggested; Nassa
olivii Bellardi, 1882 junior synonym of Nassarius semistriatus
(Brocchi, 1814); Nassa semistriata var. restitutiana
Fontannes, 1873, Nassa semistriata var. cabrierensis
Fischer & Tournouer, 1873, Nassa dertonensis Bellardi,
1882, Buccinum (Caesia) inconstans Hoernes &
Auinger, 1882, Nassa (Amyclina) dertonensis
italicistria Ruggieri, 1949, Nassa (Amyclina)
dertonensis neogigas Ruggieri, 1949 and Hinia (Hinia)
pseudocostulata Venzo & Pelosio, 1963 junior synonyms of Nassarius
striatulus striatulus (Eichwald, 1829); Buccinum
cuvierii Payaudeau, 1826 and Nassa atava Bellardi, 1882
junior synonyms of Nassarius corrugatus (Brocchi, 1814); Nassa
pinnata Bellardi, 1882, Nassa neglecta Bellardi, 1882 and Nassa
catalaunica Almera & Bofill, 1898 junior synonyms of Nassarius
elatus (Gould, 1845); Nassa crebricostulata Bellardi, 1882,
Nassa subecostata Bellardi, 1882 and Nassa nova
Bellardi, 1882 junior synonyms of Nassarius elabratus
(Doderlein,1862); Nassa pyrenaica var. compacta
Fontannes, 1879, Nassa tersa Bellardi, 1882, Nassa
heynemanni von Maltzan, 1884 and Nassa tersa var. abbreviatula
Sacco, 1904 junior synonyms of Nassarius pyrenaicus (Fontannes,
1879); Nassa semperi Bellardi, 1882 and Nassa longa
Bellardi, 1882 junior synonyms of Nassarius aldrovandii
(Bellardi, 1882); Nassa planicostata Bellardi, 1882 junior
synonym of Nassarius angulatus (Brocchi, 1814); Nassa (Hima)
rozieri Peyrot, 1925 junior synonym of Nassarius serraticosta
(Bronn, 1831); Nassa andonae Bellardi, 1882 junior synonym of Nassarius
productus (Bellardi, 1882); Buccinum carcassonni De Serres,
1829, Nassa eurosta Fontannes, 1879, Nassa d・fanconae Bellardi, 1882, Nassa forestii Bellardi,
1882, Nassa interposita Bellardi, 1882, Nassa
inaequicostata Bellardi, 1882 and Nassa fallax (Michelotti)
in Bellardi, 1882 junior synonyms of Nassarius turritus
(Borson, 1820); Eione affinis Cocconi, 1873 junior synonym of Nassarius
circumcinctus (A. Adams, 1852); Buccinum lampas Brocchi,
1814, Nassa (Desmoulaea[sic]) conglobata var. subobsoletecostata
Sacco, 1904 and Nassa (Desmoulea) conglobata altavillensis
Ruggieri et al., 1959 junior synonyms of Demoulia conglobata
(Brocchi, 1814); Nassa (Desmoulaea[sic]) conglobata var.
subobsoletecostata Sacco, 1904 junior synonym of Demoulia
pupa (Brocchi, 1814). The Italian Late Miocene and Pliocene shells
identified in the literature as Nassarius tumidus (Eichwald,
1830) are shown not to be conspecific with this Middle Miocene
Paratethyan taxon, and are considered a distinct taxon, Nassarius
tauroprimus (Sacco, 1904). The fossil and Recent populations of Nassarius
gibbosulus (Linnaeus, 1758) are considered distinct based on
morphometric analysis of their shells, the fossil shells are considered
a chronosubspecies of the Recent population, the first available name
for the fossil form being Nassarius gibbosulus pliopergibbosus
(Sacco, 1904). The Iberian assemblages fall within two groups; one with
a group of species akin to that found in the Pliocene central
Mediterranean, the north-east Spanish, Estepona Basin and Guadalquivir
Basin assemblages, and the other with a significant percentage of
distinct taxa, the Mondego Basin assemblage. Although the family is
still well represented in the Mediterranean fauna, since Pliocene times
there has been a major extinction and turnover of species. Of the 60
species recorded, only eight (13.3%) survived into the Recent faunas.
The palaeobiogeography of the Pliocene to Recent European Nassariidae
is discussed, and the strongly thermophilic character of many of the
taxa highlighted by the rapid increase in specific and generic
diversity southwards with increasing SSTs. A strong endemicity amongst
the nassariids within both Pliocene and Recent biogeographical
provinces is demonstrated, and the southward increasing number of
specific and generic taxa found in the North-East Atlantic Pliocene
biogeographical provinces follows the general pattern of Molluscan
distribution dictated by the equatorward increasing of SSTs.
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